NCHU Course Outline

Course Name	(中) 高等昆蟲分類學(8000)
Course Name	(Eng.) Advanced Insect Taxonomy
Offering Dept	Department of Entomology
Course Type	Elective	Credits	2	Teacher	Yun Hsiao
Department	Department of EntomologyPh.D	Language	English	Semester	2026-FALL
Course Description	生物系統分類學是一門歷久彌新的重要學門，其起源自基於形態特徵觀察的分類鑑定，以及隨之而生的物種命名發表。然而，當代的系統分類學研究不僅止於此。隨著時代演進，除了高解析度觀察設備與顯微攝影技術的突飛猛進，更因為標準化分子親緣分析、物種界定、地理分布與環境因子圖層分析，以及親緣比較方法（PCMs）的成熟，使其在研究深度與廣度上有了飛躍式的進展。本課程將橫跨親緣分析、物種界定、空間分析、幾何形態學、量化形態分析以及親緣比較分析法，介紹其實用分析方法並帶領學生動手實作。修課學生應具備基本的親緣分析、統計學及 R 語言操作能力。 Biological systematics is an enduring and vital discipline, originating from morphology-based species identification and subsequent taxonomic descriptions. However, contemporary systematic research extends far beyond these traditional foundations. Driven by rapid advancements in high-resolution imaging technologies, alongside the integration of standardized molecular phylogenetics, species delimitation, spatial analysis, and phylogenetic comparative methods (PCMs), the field has experienced a quantum leap in both its scope and depth. This course spans phylogenetic reconstruction, species delimitation, spatial analysis, geometric morphometrics, quantitative morphological analysis, and evolutionary comparative methods. Blending theoretical foundations with hands-on practical sessions, students are expected to possess baseline knowledge in phylogenetics, statistics, and R programming.
Prerequisites				self-directed learning in the course	Y

Relevance of Course Objectives and Core Learning Outcomes(%)

Teaching and Assessment Methods for Course Objectives

Course Objectives

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Teaching Methods

Assessment Methods

1. 了解符合當代前沿標準的系統分類學研究分析方法。
To understand and master cutting-edge analytical methods that meet the rigorous standards of contemporary systematic research.
2. 充實並拓展個人研究專題的廣度與深度。
To enrich and expand both the breadth and depth of students' individual research projects.

topic Discussion/Production

Exercises

Lecturing

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Oral Presentation

Course Content and Homework/Schedule/Tests Schedule

Week	Course Content
Week 1	9/10:課程概述 (Introduction to the Course) & 何謂現代標準的系統分類學研究? (What Constitutes a Modern Standard Systematic Study?)
Week 2	9/17:分子親緣分析：序列比對與資料分區、分子演化模型選擇、拔靴法、與鄰接聚類法 (Molecular Phylogenetic Analysis: Alignment and Partitions, Evolutionary Model Selection, Bootstrapping and Neighbor-Joining Method)
Week 3	9/24:分子親緣分析：最大似然法與貝氏親緣分析 (Molecular Phylogenetic Analysis: Maximum Likelihood and Bayesian Methods)
Week 4	10/1:分子物種界定 (Molecular Species Delimitation)
Week 5	10/8:幾何形態分析 (Morphometrics)
Week 6	10/15:物種分布地圖和空間分析 (Species Distribution Map and Spatial Analysis)
Week 7	10/22:引導式自主期中專題研究與方法學諮詢 (Guided Independent Midterm Project Research and Methodology Consultation)
Week 8	10/29:期中口頭報告 (Mid-Term Oral Presentation)
Week 9	11/5:利用離散特徵進行親緣關係重建 (Morphology-based Phylogenetic Analysis)
Week 10	11/12:形態多樣性分析 (Morphological Diversity Analysis)
Week 11	11/19:分化時間估算 (Divergence Time Estimation)
Week 12	11/26:生物地理歷史重建 (Ancestral Distribution Reconstruction)
Week 13	12/3:祖徵估算 (Ancestral Character State Reconstruction)
Week 14	12/10: 親緣廣義線性混合模型 (Phylogenetic Generalized Linear Mixed Model)
Week 15	12/17:引導式自主期末專題研究與方法學諮詢 (Guided Independent Final Project Research and Methodology Consultation)
Week 16	12/24:期末口頭報告 (Final Oral Presentation)
self-directed learning	03.Preparing presentations or reports related to industry and academia. 將要求學生利用課堂學習知識，自行選擇主題、分析資料以進行兩次口頭報告，自主學習內容達2學分4小時之要求。 To fulfill the 2-credit, 4-hour independent learning requirement, students will independently select research topics, analyze data, and deliver two oral presentations by applying the knowledge learned in class.

Evaluation

期中口頭報告 (40%); 期末口頭報告 (40%); 出席率 (20%)
Mid-Term Oral Presentation (40%); Final Oral Presentation (40%); Presence (20%)
##出席規定 Presence
曠課
1) 按學校規定：學生缺課時數達該全學期上課時數三分之一者，不得參加該科目之學期考試，該科目學期成績以零分計
2) 若未超過三分之一者，每次缺課扣學期總分三分，扣至分數佔比20％為止
Unexcused Absences:
1) University Regulations: In accordance with school policy, any student whose total hours of absence reach one-third of the total scheduled class hours for the semester shall be disqualified from taking the final examination. In such cases, the final grade for the course will be recorded as zero.
2) Grading Deductions: For students whose absences do not exceed the one-third threshold, a 3-point deduction will be applied to the final grade for each occurrence, up to a maximum total deduction of 20% of the final grade.

Textbook & other References

1. Brower, A.V.Z. and Schuh, R.T. 2021. Biological Systematics: Principles and Applications (3rd edn). Ithaca, NY; online edn, Cornell Scholarship Online. https://doi.org/10.7591/cornell/9781501752773.001.0001
2. Bromham, L. 2016. An Introduction to Molecular Evolution and Phylogenetics (2nd edn). Oxford University Press.
3. Compiled from selected academic journal papers參考學術期刊論文自行彙編

Teaching Aids & Teacher's Website

I-Learning

Office Hours

by email (教師電子郵件約時間)

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include experience courses：N

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Update Date, year/month/day：2026/06/16 14:14:36	Printed Date, year/month/day：2026 / 6 / 19
The second-hand book website：http://www.myub.com.tw/